Botany 201 - Taxonomy of Flowering Plants - Spring, 1995

Lecture Examination II

Instructions: This is a replicate 'key' of the second lecture exam. The final exam will be partially comprehensive (100 points). It is therefore important to know correct responses to questions of the three hour exams. Correct responses are highlighted for single response questions. These are internal links that, on selection, will take you to an overview discussion of the question and responses. Selection of the highlighted correct response in the overview, or use of the 'back' button on your browser, will take you back to the question. Similar linkage for 'matching' questions is via the verb of the question.

1. The following sequence of taxonomic names represents which sequence of ranks in the taxonomic hierarchy: Magnoliales, Magnoliaceae, Magnoliopsida, Magnoliidae

         a.  Genus, Order, Family, Subclass
         b.  Order, Family, Class, Subclass
         c.  Genus, Family, Order, Subclass
         d.  Family, Subclass, Order, Division
         e.  Order, Division, Class, Subclass
  • Identify the Family: (questions 2 - 8)

  • 2. Woody, syncarpous plants with simple leaves and unisexual flowers:

            a.  Fagaceae    b. Magnoliaceae    c. Juglandaceae  
                  d. Winteraceae    e. Caryophyllaceae
    3. Woody, apetalous plants that often produce multiple fruits:
                 a. Magnoliaceae    b. Moraceae    c. Fagaceae  
                      d. Juglandaceae    e. Urticaceae
    4. Woody, syncarpous plants with compound leaves and unisexual flowers:
            a. Amaranthaceae    b.  Cannabaceae    c. Juglandaceae  
                          d. Papaveraceae    e. Moraceae
    5. Mostly herbs with swollen nodes, alternate leaves, and perfect flowers:
          a. Papaveraceae    b. Polygonaceae    c. Caryophyllaceae 
                        d. Cactaceae    e. Moraceae
    6. Dioecious, anemophilous, herbaceous plants that are often placed within the Moraceae:
             a.  Juglandaceae    b. Cannabaceae    c. Papaveraceae   
                       d. Phytolaccaceae    e. Cactaceae
    7. Fruit often a denticidal capsule:
               a. Papaveraceae   b. Cactaceae    c. Winteraceae  
                    d.  Caryophyllaceae   e. Cannabaceae
    8. Both herbaceous and woody plants with betalains, full syncarpy, and alternate leaves:
         a. Papaveraceae    b.  Chenopodiaceae    c. Ranunculaceae  
                 d.  Phytolaccaceae    e. Caryophyllaceae
  • Link the appropriate Subclass to key characters: (questions 9 - 11)
  •      a. Magnoliidae    b.  Hamamelidae    c. Caryophyllidae
    9. Often dominant elements of the world's deciduous forests, reproductive system often based on anemophily.

     10. Mostly herbs that often show 'beaked' seeds which reflect a peripheral embryo position.

     11. Mostly woody and apocarpous.

  • Match term with definition: (questions 12 - 15)

  • 12. halophyte a. diploid nutritive tissue 13. perisperm b. pollinated by insects 14. anthesis c. flower open and fully expanded 15. entomophilous d. wind pollinated e. plant adapted to saline conditions

  • Associate economically important genera to their Family: (questions 16 - 20)

  •  

     

    16.  Ficus        a.  Moraceae
    17.  Beta         b.  Polygonaceae
    18.  Fagopyrum    c.  Chenopodiaceae
    19.  Liriodendron d.  Magnoliaceae
    20.  Persea       e.  Lauraceae
    21. The nomenclatural type that was designated by the original author as the type specimen is known as the:
                 a. isotype    b. neotype    c. paratype
                       d. holotype    e. lectotype
    22. Select the family characterized by latex production:
           a.  Juglandaceae   b.  Magnoliaceae    c.  Papaveraceae 
                         d. Winteraceae    e. Fagaceae
    23. The __________ matures to form a fruit, whereas a mature _______ is called a seed.
       a. egg, ovule    b. carpel, pericarp   c. ovary, microgametophyte
                     d. funiculus, ovary    e. ovary, ovule
    24. Select the family characterized by opposite leaves:
         a. Magnoliaceae    b.  Juglandaceae    c.  Caryophyllaceae
                         d. Papaveraceae    e. Fagaceae
    25. Select the xerophytic taxon:
               a. Opuntia   b. Magnoliaceae     c.  Fagaceae  
                        d. Dianthus    e. Papaver
    26. Which taxon stands as a possible link between the Magnoliopsida and the Liliopsida?:
               a. Magnoliaceae    b.  Thalictrum    c.  Nymphaeales  
                          d. Caryophyllales    e. Lauraceae
    27. Which of the following is unique to the Magnoliophyta:
       a. pericarp    b. xylem    c. funiculus    d. meristem    e. ovule
    28. Who is responsible for bringing the Latin binomial into common usage as an essential part of plant nomenclature?
         a.   A. P. de Candolle     b.  A. Engler       c.   J. Lamark    
                     d.   C. Bessey     e.   C. Linnaeus
  • Identify the Family: (questions 29 - 34)

  • 29. Apocarpous, woody plants with perfect flowers and a circular stipule scar:

            a. Magnoliaceae    b. Fagaceae    c.  Juglandaceae 
                      d. Moraceae    e. Urticaceae
    30. Woody perennials of the southern hemisphere that lack vessel elements in the xylem:
           a. Magnoliaceae    b.  Winteraceae    c. Papaveraceae 
                     d. Fagaceae    e. Juglandaceae
    31. Epigynous, often succulent, herbs with fugacious leaves:
           a.  Cactaceae    b.  Urticaceae    c.  Winteraceae  
                d. Amaranthaceae    e. Caryophyllaceae
    32. Associated with the Chenopodiaceae, but differing by a lack of connation in the perianth:
           a.  Moraceae   b. Papaveraceae    c. Polygonaceae 
                d. Amaranthaceae   e. Caryophyllaceae
    33. Apocarpous herbs of the Magnoliidae:
        a. Papaveraceae     b.  Lauraceae    c.  Chenopodiaceae  
                   d. Cactaceae    e. Ranunculaceae
    34. Syncarpous herbs of the Hamamelidae:
           a.  Urticaceae    b. Polygonaceae    c. Moraceae  
                  d. Lauraceae    e.  Fagaceae
  • Match structures with associated taxa: (questions 35 - 39)
  • 35.  areole                 a.  Caryophyllaceae
    36.  synconium              b.  Ficus
    37.  ocrea                  c.  Polygonaceae
    38.  poricidal capsule      d.  Papaveraceae
    39.  clawed petals          e.  Cactaceae
    40. We did not have an International Code of Botanical Nomenclature until ______, but the Principal of Priority extends back in time to _____.
          a.  1930, 1753    b. 1856, 1930    c. 1753, 1960    
                   d. 1903, 1735    e. 1830, 1915

    Response Overviews 

    1. The following sequence of taxonomic names:

    Magnoliales, Magnoliaceae, Magnoliopsida, Magnoliidae

     represents which sequence of ranks in the taxonomic hierarchy: (or, 'Can you link nomenclature (name endings) with rank?')
     
     

  • a. Genus, Order, Family, Subclass
  • b. Order, Family, Class, Subclass - APPROACH: Note rank of 1st name on list and screen available responses for those proper rank (order - '-ales' in this case) in the first position - only 2 (b. and e.) in this case. Note rank of next name in the sequence and check the two possibilities - only one in this case (b.) with the 'order-family' sequence.
  • c. Genus, Family, Order, Subclass
  • d. Family, Subclass, Order, Division
  • e. Order, Division, Class, Subclass

  • 2. Woody, syncarpous plants with simple leaves and unisexual flowers:
     
     

  • a. Fagaceae - all characters fit this family of the Hamameliidae
  • b. Magnoliaceae - woody with simple leaves, but apocarpous and perfect flowers
  • c. Juglandaceae - all characters fit BUT genera of this family show compound leaves
  • d. Winteraceae - woody with simple leaves, but apocarpous and perfect flowers, as sister family (response option 'b') of the Magnoliidae
  • e. Caryophyllaceae - matches with syncarpy and simple leaves, but elements of this family, all herbaceous, usually show perfect flowers.

  • 3. Woody, apetalous plants that often produce multiple fruits:
     
     

  • a. Magnoliaceae - Woody, but polypetalous (separate petals in the corolla) and no multiple fruits (although aggregate fruiting structures often develop from apocarpous gynoecia).
  • b. Moraceae - fits all characters
  • c. Fagaceae - Woody and often apetalous flowers in a catkin or 'ament', but fruit is typically a nut
  • d. Juglandaceae - as response 'c' above
  • e. Urticaceae - apetalous, but not woody and no multiple fruits

  • 4. Woody, syncarpous plants with compound leaves and unisexual flowers:
     
     

  • a. Amaranthaceae - not woody and with simple leaves
  • b. Cannabaceae - also not woody
  • c. Juglandaceae - fits all characters
  • d. Papaveraceae - not woody with perfect flowers
  • e. Moraceae - fits all characters except compound leaves

  • 5. Mostly herbs with swollen nodes, alternate leaves, and perfect flowers:
     
     

  • a. Papaveraceae - fits all characters EXCEPT 'swollen nodes'
  • b. Polygonaceae - fits all characters
  • c. Caryophyllaceae - fits all characters EXCEPT alternate leaves
  • d. Cactaceae - as response 'a'
  • e. Moraceae - woody, flowers not perfect and NO swollen nodes

  • 6. Dioecious, anemophilous, herbaceous plants that are often placed within the Moraceae:
     
     

  • a. Juglandaceae - woody, monoecious and NOT associated with the Moraceae or, for that matter, the Urticales
  • b. Cannabaceae - fits all characters
  • c. Papaveraceae - perfect-flowered, entomophilous element of the Magnoliidae
  • d. Phytolaccaceae - perfect-flowered, entomophilous element of the Caryophyllidae
  • e. Cactaceae - perfect-flowered, entomophilous element of the Caryophyllidae

  • 7. Fruit often a denticidal capsule:
     
     

  • a. Papaveraceae - fruit a capsule, but often poricidal
  • b. Cactaceae - fruit a berry
  • c. Winteraceae - fruit unicarpellate, a follicle
  • d. Caryophyllaceae - this - dehiscence by terminal 'valves' is one of many good key characters for this family
  • e. Cannabaceae - fruit single-seeded - an achene

  • 8. Both herbaceous and woody plants with betalains, full syncarpy, and alternate leaves: (The character 'betalains' points toward a subclass (Caryophyllidae) - other characters allow 'sorting' of families circumscribed within that subclass)
     
     

  • a. Papaveraceae - all herbs and no betalains
  • b. Chenopodiaceae - all characters are consistent with this family
  • c. Ranunculaceae - all apocarpous herbs with no betalains
  • d. Phytolaccaceae - betalains, but partial syncarpy
  • e. Caryophyllaceae - betalains, syncarpy, but opposite leaves

  • Link the appropriate Subclass to key characters: (questions 9 - 11)

  •  9. Often dominant elements of the world's deciduous forests, reproductive system often based on anemophily: b. Hamamelidae

     10. Mostly herbs that often show 'beaked' seeds which reflect a peripheral embryo position: c. Caryophyllidae

     11. Mostly woody and apocarpous: a. Magnoliidae


  • Match term with definition: (questions 12 - 15)
  • 12. halophyte: e. plant adapted to saline conditions
  • 13. perisperm:a. diploid nutritive tissue
  • 14. anthesis:c. flower open and fully expanded
  • 15. entomophilous:b. pollinated by insects

  • (response 'd. wind pollinated' = anemophilous)

  • Associate economically important genera to their Family: (questions 16 - 20)

  •  

     

  • 16. Ficus (fig): a. Moraceae
  • 17. Beta (beet): c. Chenopodiaceae
  • 18. Fagopyrum (buckwheat): b. Polygonaceae
  • 19. Liriodendron: (yellow poplar): d. Magnoliaceae
  • 20. Persea (avocado): e. Lauraceae


  • 21. The nomenclatural type that was designated by the original author as the type specimen is known as the:
     
     

  • a. isotype - a duplicate of the original (holotype) type specimen
  • b. neotype - a specimen selected when all original specimens and their duplicates have been destroyed
  • c. paratype - a specimen, other than the holotype or isotype, that was cited in the original publication of the name
  • d. holotype - the one specimen used or designated by the author in the original publication as the nomenclatural type
  • e. lectotype - specimen chosen by a later worker from original material studied by the author of the species name

  • 22. Select the family characterized by latex production:

  • a. Juglandaceae
  • b. Magnoliaceae
  • c. Papaveraceae - latex production, like opposite leaves, is a 'subset' character. Only two families covered to date show latex production; the Papaveraceae and Moraceae.
  • d. Winteraceae
  • e. Fagaceae

  • 23. The __________ matures to form a fruit, whereas a mature _______ is called a seed.
     
     

  • a. egg, ovule
  • b. carpel, pericarp
  • c. ovary, microgametophyte
  • d. funiculus, ovary
  • e. ovary, ovule - basic terms that are fundamental to understanding the gynoecium

  • 24. Select the family characterized by opposite leaves:

  • a. Magnoliaceae
  • b. Juglandaceae
  • c. Caryophyllaceae -opposite leaves, like latex production, is a 'subset' character. This is the only family covered to date show opposite leaf arrangement as a circumscribing character.
  • d. Papaveraceae
  • e. Fagaceae

  • 25. Select the xerophytic (adapted to DRY conditions) taxon:

  • a. Opuntia - a xerophytic genus that is part of a xerophytic family
  • b. Magnoliaceae
  • c. Fagaceae
  • d. Dianthus
  • e. Papaver

  • 26. Which taxon stands as a possible link between the Magnoliopsida and the Liliopsida?:

  • a. Magnoliaceae - a basal element, but no linkage to the monocots
  • b. Thalictrum - relatively specialized genus (anemophilous) of a specialized family (Ranunculaceae - herbaceous)
  • c. Nymphaeales - classic (fide Charles Bessy) archaic flower BUT herbaceous and without vessels (possibly lost due to aquatic adaptations) see Cronquist, A. 1988. The Evolution and Classification of Flowering Plants (NYBG). pp. 451-457.
  • d. Caryophyllales - few archaic features (reduced numbers of floral appendages, syncarpy, synsepaly) and therefore not a likely candidate.
  • e. Lauraceae - basal at the subclass level (Magnoliidae), but, within this context, relatively specialized (reduced number of floral appendages).

  • 27. Which of the following is unique to the Magnoliophyta(=flowering plants):

  • a. pericarp - ovary wall; ONLY the flowering plants (Magnoliophyta) have an ovary.
  • b. xylem - conductive tissue that is present in ALL vascular plants (pteriodphytes [ferns], gymnosperms AND angiosperms)
  • c. funiculus - stalk of the ovule, present on all seed plants (gymnosperms AND angiosperms)
  • d. meristem - centers of cell division that tend of define the Plant Kingdom
  • e. ovule - see response 'c'

  • 28. Who is responsible for bringing the Latin binomial into common usage as an essential part of plant nomenclature?

  • a. A. P. de Candolle - played a part in development of the ICBN
  • b. A. Engler - responsible for the first major classification system that carried a phylogenetic rationale
  • c. J. Lamark - known for non-Mendelian notions of inheritance
  • d. C. Bessey - established principals of phylogenetic connections that are employed in modern systems of classification
  • e. C. Linnaeus - first to use the binomial in a major work - Species Plantarum, 1753.

  • 29. Apocarpous, woody plants with perfect flowers and a circular stipule scar:

  • a. Magnoliaceae - all characters apply
  • b. Fagaceae - woody, but syncarpous and flowers unisexual
  • c. Juglandaceae - as response 'b'
  • d. Moraceae - circular stipule scar, but flowers imperfect
  • e. Urticaceae - herbaceous with unisexual flowers

  • 30. Woody perennials of the southern hemisphere that lack vessel elements in the xylem:

  • a. Magnoliaceae Woody perennials with a north-temperate distribution and vessels
  • b. Winteraceae - often placed as one of the most archaic elements of the basal dicot subclass Magnoliidae
  • c. Papaveraceae - an herbaceous family with vessels and a worldwide distribution
  • d. Fagaceae - Woody plants, but with a wide distribution and, as is the case with all Hamamelidae, vessel elements in the xylem.
  • e. Juglandaceae - Woody plants, but with a north-temperate and, as is the case with all Hamamelidae, vessel elements in the xylem.

  • 31. Epigynous, often succulent, herbs with fugacious leaves:

  • a. Cactaceae - Of possible responses, the only epigynous family
  • b. Urticaceae - not succulent with functional leaves
  • c. Winteraceae - not herbaceous
  • d. Amaranthaceae - flower hypogynous, leaves photosynthetic, usually not succulent
  • e. Caryophyllaceae - as response 'd'.

  • 32. Associated with the Chenopodiaceae, but differing by a lack of connation in the perianth: (association with the Chenopodiaceae points toward a family of the Caryophyllidae)

  • a. Moraceae - Hamamelidae
  • b. Papaveraceae - Magnoliidae
  • c. Polygonaceae - Caryophyllidae, but Polygonales
  • d. Amaranthaceae - a visit to the class assignment when we covered the chenopods, i.e., note differences between Chenopodiaceae and its allied family.
  • e. Caryophyllaceae - Caryophyllidae, Caryophyllales - but not the best response (genera typically show connation in the perianth)

  • 33. Apocarpous herbs of the Magnoliidae:

  • a. Papaveraceae - Magnoliidae, but syncarpous
  • b. Lauraceae - Magnoliidae, but woody and syncarpous
  • c. Chenopodiaceae - Caryophyllidae
  • d. Cactaceae - Caryophyllidae
  • e. Ranunculaceae - only response that fits the characters - the first POP QUIZ revisited

  • 34. Syncarpous herbs of the Hamamelidae:

  • a. Urticaceae - essentially ALL elements of the Hamamelidae are syncarpous, but most are woody. 'Nettles' are, however, herbaceous plants
  • b. Polygonaceae - Caryophyllidae
  • c. Moraceae - Hamamelidae, but woody
  • d. Lauraceae - Magnoliidae
  • e. Fagaceae - Hamamelidae, but woody

  • Match structures with associated taxa: (questions 35 - 39)

  • 35. areole - nodal spine cluster resulting from modification of the lateral bud in the: e. Cactaceae
  • 36. synconium - the strange, enclosed multiple fruit ('fig') that is unique to: b. Ficus of the Moraceae
  • 37. ocrea - stipular sheath that is a good key character for many genera of the: c. Polygonaceae
  • 38. poricidal capsule - type of dehiscence that is rare in the angiosperms but quite common among elements of the: d. Papaveraceae
  • 39. clawed petals - a distinctive feature that marks many genera of the: a. Caryophyllaceae 

  • 40. We did not have an International Code of Botanical Nomenclature until 1930 (final merger of European and American 'codes'), but the Principal of Priority extends back in time to 1753 (publication of Species Plantarum).


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