Taxonomy of Flowering Plants - LECTURE NOTES - Spring, 1998
Hugh D. Wilson - rm. 306, Butler Hall
23 February 1998
The Caryophyllidae
Family Overview - The Caryophyllales

Chenopodiaceae - the Goosefoot Family

Diversity:  A taxonomically difficult group of 102 genera and about 1,400 species.  These are mostly herbs, with many weedy annuals, but also some fairly common fruticose taxa.  Many 'chenopods' are halophytic plants adapted to soils with a high salt content, such as beaches and dry lakes.  Since water retention is a primary problem in this type of environment, many species show a fleshy succulence.  The ethnoflora includes the beet (Beta vulgaris), spinach (Spinacia oleracea) and the 'pseudoceral' quinoa (Chenopodium quinoa).

Distribution:  World wide, especially xeric or saline habitats with centers of diversity in South America and Australia.  The Texas flora includes 16 genera and 60 species, many of these common weeds of agricultural fields.

Floral structure:

 

Significant features:  A difficult family in that the flowers are small and the plants more or less 'streamlined' with few conspicuous characters.  Elements of the family are, however, often common and ecologically important.  Field recognition keys on an 'eye' focused on reduced flowers, often perfect but sometimes unisexual, producing the distinctive, often uniovulate 'beaked' fruit depicted below and a 'fleshy' or succulent aspect to the plant.  Epigyny is rare in the family (only Beta).

   Overview of a typical beet plant (Beta vulgaris) from Kohler's Medicinal Plants
 
Chenopodium berlandieri with sectioned fruit (left - pericarp, testa, perisperm, embryo), whole fruit (center - reticulate pericarp, darker testa, and 'beak' of the radical), and (right)flower at anthesis:
 
 

 
 
 
 

   Chenopodium  missouriense  - inflorescence - at anthesis

 
The Amaranthaceae (pigweed family) is a 'sister' group to the Chenopodiaceae in that all working with classification of the Caryophyllales, regardless of approach, agree that the two families are closely related.  They also share similarities in size, structure, ecological amplitude, and distribution.  Check your text, or information available on the network, to determine how one might distinguish between these two families if only the flower was available for inspection.

More information on the Chenopodiaceae, including species diversity in North America and California


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