Verbenaceae - the Vervain or Verbena Family
Distribution: Worldwide, but as opposed to other elements of the Lamiales, with greatest diversity in the tropics. We have 14 genera and 69 species in the Texas flora, including a single endemic species in South Texas.
Floral structure:


Significant features:
This family carries the distinctive gynoecium of the Lamiales
in that the ovary usually shows a second septum, producing 4
locules
and each of these has only a single seed. However, the lobing in
most taxa is not as extensive as that found in the Boraginaceae and
Lamiaceae
and the style is not gynobasic, it is usually positioned at the
tip
of the ovary. Taxa of the Verbenaceae, especially the
herbs,
tend of show quadrangular stems
and
all have opposite leaves.
This
similarity to the Lamiaceae also extends to corolla
zygomorphy but this is less developed in the Verbenaceae and
usually expressed as an asymmetry in the corolla lobes as opposed to
the
bilabiate ('two-lipped') corolla configuration common among the mints..
Verbena:
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Callicarpa americana:
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More information on the Verbenaceae